The spines macro allows an arbitrary object to be placed at intervals along a spline path.
Because spines is primarily meant to complement the tentacle macro it uses many of the same inputs as the tentacle to enable the two to match up easily.
In addition to the tentacle inputs spines also takes a width macro in addition to the radius macro which allows the object to be scaled independently of the radius of the tentacle. It also contains the option to scale the object in proportion to the radius macro as well.